:root {
  --ink: #17252d;
  --muted: #53656e;
  --line: #d8e0e3;
  --surface: #ffffff;
  --surface-muted: #f2f6f7;
  --ocean: #075b71;
  --ocean-dark: #064455;
  --ocean-pale: #dceff3;
  --status: #f3e7c8;
  --status-border: #d8bd77;
  --danger: #a42a2a;
  --danger-pale: #fff0f0;
  --shadow: 0 10px 30px rgb(21 49 59 / 9%);
  --content-width: 1500px;
  color-scheme: light;
  font-family: system-ui, -apple-system, BlinkMacSystemFont, "Segoe UI", sans-serif;
  font-synthesis: none;
}

*,
*::before,
*::after {
  box-sizing: border-box;
}

html {
  scroll-behavior: smooth;
}

body {
  margin: 0;
  background: var(--surface-muted);
  color: var(--ink);
  line-height: 1.55;
}

a {
  color: var(--ocean);
  text-underline-offset: 0.18em;
}

a:hover {
  color: var(--ocean-dark);
}

button,
select,
input {
  font: inherit;
}

button:focus-visible,
select:focus-visible,
input:focus-visible,
a:focus-visible,
[tabindex="-1"]:focus-visible {
  outline: 3px solid #ee9b36;
  outline-offset: 3px;
}

.skip-link {
  position: fixed;
  z-index: 20;
  top: 0.75rem;
  left: 0.75rem;
  padding: 0.65rem 0.9rem;
  border-radius: 0.35rem;
  background: var(--ink);
  color: #fff;
  transform: translateY(-160%);
}

.skip-link:focus {
  transform: translateY(0);
}

.site-header {
  border-bottom: 1px solid rgb(255 255 255 / 18%);
  background:
    radial-gradient(circle at 82% 20%, rgb(85 181 194 / 28%), transparent 28rem),
    linear-gradient(125deg, #063e50, #087085);
  color: #fff;
}

.site-header__inner,
.species-toolbar__inner,
main,
.site-footer {
  width: min(calc(100% - 2rem), var(--content-width));
  margin-inline: auto;
}

.site-header__inner {
  padding: clamp(2.5rem, 7vw, 5.5rem) 0 clamp(2.25rem, 5vw, 4rem);
}

.eyebrow {
  margin: 0 0 0.35rem;
  color: inherit;
  font-size: 0.76rem;
  font-weight: 750;
  letter-spacing: 0.12em;
  text-transform: uppercase;
}

.site-header h1 {
  max-width: 18ch;
  margin: 0;
  font-size: clamp(2rem, 5vw, 4.5rem);
  font-weight: 720;
  letter-spacing: -0.045em;
  line-height: 0.98;
}

.species-toolbar {
  position: sticky;
  z-index: 10;
  top: 0;
  border-bottom: 1px solid var(--line);
  background: rgb(255 255 255 / 96%);
  box-shadow: 0 4px 16px rgb(21 49 59 / 7%);
  backdrop-filter: blur(10px);
}

.species-toolbar__inner {
  display: flex;
  align-items: end;
  justify-content: space-between;
  gap: 1rem;
  padding-block: 0.85rem;
}

.species-picker {
  flex: 1 1 30rem;
  max-width: 44rem;
}

.species-picker label {
  display: block;
  margin-bottom: 0.25rem;
  color: var(--muted);
  font-size: 0.78rem;
  font-weight: 700;
  letter-spacing: 0.05em;
  text-transform: uppercase;
}

.species-combobox {
  position: relative;
}

.species-search {
  width: 100%;
  min-height: 2.75rem;
  padding: 0.45rem 0.7rem;
  border: 1px solid #a8b8be;
  border-radius: 0.4rem;
  background: var(--surface);
  color: var(--ink);
}

.species-search::placeholder {
  color: var(--muted);
}

.species-options {
  position: absolute;
  z-index: 30;
  top: calc(100% + 0.35rem);
  width: 100%;
  max-height: min(50vh, 24rem);
  margin: 0;
  padding: 0.3rem;
  overflow-y: auto;
  border: 1px solid #a8b8be;
  border-radius: 0.45rem;
  background: var(--surface);
  box-shadow: var(--shadow);
  list-style: none;
}

.species-option,
.species-no-results {
  padding: 0.55rem 0.65rem;
  border-radius: 0.3rem;
}

.species-option {
  cursor: pointer;
}

.species-option:hover,
.species-option.is-active {
  background: var(--ocean-pale);
  color: var(--ocean-dark);
}

.species-no-results {
  color: var(--muted);
  font-style: italic;
}

.species-nav {
  display: flex;
  flex: 0 0 auto;
  gap: 0.45rem;
}

.species-nav button {
  min-height: 2.75rem;
  padding: 0.45rem 0.8rem;
  border: 1px solid #a8b8be;
  border-radius: 0.4rem;
  background: var(--surface);
  color: var(--ocean-dark);
  cursor: pointer;
  font-weight: 650;
}

.species-nav button:hover:not(:disabled) {
  border-color: var(--ocean);
  background: var(--ocean-pale);
}

.species-nav button:disabled {
  cursor: not-allowed;
  opacity: 0.45;
}

main {
  min-height: 55vh;
  padding-block: clamp(1.5rem, 4vw, 3.5rem) 4rem;
}

.app-message {
  width: min(100%, 50rem);
  margin: 0 auto 1.5rem;
  padding: 0.85rem 1rem;
  border: 1px solid #a6cbd3;
  border-radius: 0.45rem;
  background: var(--ocean-pale);
  color: var(--ocean-dark);
  text-align: center;
}

.app-message--error {
  border-color: #e0a1a1;
  background: var(--danger-pale);
  color: var(--danger);
}

.species-header,
.metadata-panel,
.text-section {
  width: min(100%, 72rem);
  margin-inline: auto;
}

.species-header {
  display: flex;
  align-items: end;
  justify-content: space-between;
  gap: 2rem;
  padding: clamp(1.25rem, 3vw, 2rem);
  border-radius: 0.65rem 0.65rem 0 0;
  background: var(--ocean-dark);
  color: #fff;
}

.species-header h2 {
  margin: 0;
  font-size: clamp(2rem, 5vw, 3.75rem);
  letter-spacing: -0.035em;
  line-height: 1.03;
}

.scientific-name {
  margin: 0.55rem 0 0;
  color: rgb(255 255 255 / 78%);
  font-family: Georgia, "Times New Roman", serif;
  font-size: clamp(1.1rem, 2vw, 1.35rem);
  font-style: italic;
}

.status-badges {
  display: flex;
  flex: 0 1 29rem;
  flex-wrap: wrap;
  justify-content: flex-end;
  gap: 0.5rem;
}

.status-badge {
  padding: 0.45rem 0.65rem;
  border: 1px solid var(--status-border);
  border-radius: 0.4rem;
  background: var(--status);
  color: #463812;
  font-size: 0.88rem;
}

.status-badge strong {
  display: block;
  font-size: 0.7rem;
  letter-spacing: 0.06em;
  text-transform: uppercase;
}

.metadata-panel {
  display: flex;
  flex-wrap: wrap;
  align-items: center;
  justify-content: space-between;
  gap: 1.5rem;
  padding: 1.15rem clamp(1.25rem, 3vw, 2rem);
  border: 1px solid var(--line);
  border-top: 0;
  border-radius: 0 0 0.65rem 0.65rem;
  background: var(--surface);
  box-shadow: var(--shadow);
}

.taxonomy-list {
  display: flex;
  flex-wrap: wrap;
  gap: 1rem 2.5rem;
  margin: 0;
}

.taxonomy-list div {
  display: grid;
  gap: 0.05rem;
}

.taxonomy-list dt {
  color: var(--muted);
  font-size: 0.73rem;
  font-weight: 700;
  letter-spacing: 0.07em;
  text-transform: uppercase;
}

.taxonomy-list dd {
  margin: 0;
  font-weight: 650;
}

.external-links {
  display: flex;
  flex-wrap: wrap;
  justify-content: flex-end;
  gap: 0.5rem 1rem;
}

.external-links a::after {
  content: " ↗";
  font-size: 0.8em;
}

.reference-links {
  display: flex;
  flex: 1 1 100%;
  flex-wrap: wrap;
  justify-content: flex-end;
  gap: 0.35rem 0.85rem;
  border-top: 1px solid var(--line);
  padding-top: 0.75rem;
  font-size: 0.88rem;
}

.reference-links::before {
  flex: 1 1 100%;
  color: var(--muted);
  content: "Reports and references";
  font-size: 0.7rem;
  font-weight: 700;
  letter-spacing: 0.07em;
  text-align: right;
  text-transform: uppercase;
}

.reference-links a::after {
  content: " ↗";
  font-size: 0.8em;
}

.text-section {
  margin-top: 1.25rem;
  padding: 1.25rem clamp(1.25rem, 3vw, 2rem);
  border: 1px solid var(--line);
  border-radius: 0.65rem;
  background: var(--surface);
}

.text-section h3 {
  margin: 0 0 0.65rem;
  font-size: 1rem;
}

.text-section ul {
  margin: 0;
  padding-left: 1.25rem;
}

.text-section li + li,
#notes-list p + p {
  margin-top: 0.5rem;
}

.reference-heading {
  overflow-wrap: anywhere;
}

.reference-label {
  font-weight: 700;
}

.reference-link {
  font-weight: 650;
}

.text-section details {
  margin-top: 0.15rem;
  color: var(--muted);
  font-size: 0.92rem;
}

.text-section summary {
  color: var(--ocean);
  cursor: pointer;
  font-size: 0.83rem;
}

.reference-text {
  display: block;
  margin-top: 0.2rem;
  overflow-wrap: anywhere;
  color: var(--muted);
}

#notes-list p {
  margin: 0;
  white-space: pre-line;
}

.figures-section {
  margin-top: clamp(2.5rem, 7vw, 5.5rem);
}

.section-heading {
  display: flex;
  align-items: end;
  justify-content: space-between;
  gap: 1.5rem;
  width: min(100%, 72rem);
  margin: 0 auto 1rem;
}

.section-heading .eyebrow {
  color: var(--ocean);
}

.section-heading h3 {
  margin: 0;
  font-size: clamp(1.6rem, 3vw, 2.25rem);
}

.section-heading > p {
  margin: 0;
  color: var(--muted);
  font-size: 0.9rem;
}

.figures {
  display: grid;
  gap: clamp(1.25rem, 3vw, 2.5rem);
}

.synopsis-figure {
  margin: 0;
}

.figure-frame {
  position: relative;
  min-height: 14rem;
  overflow: hidden;
  border: 1px solid var(--line);
  border-radius: 0.5rem;
  background:
    linear-gradient(110deg, transparent 25%, rgb(255 255 255 / 70%) 40%, transparent 55%) 0 0 / 220% 100%,
    #e4eaec;
  box-shadow: var(--shadow);
}

.figure-frame.is-loading {
  animation: loading-sheen 1.8s linear infinite;
}

.figure-frame img {
  display: block;
  width: 100%;
  height: auto;
  background: #fff;
}

.figure-loading,
.figure-error {
  position: absolute;
  inset: 1rem auto auto 50%;
  z-index: 1;
  margin: 0;
  padding: 0.4rem 0.65rem;
  border-radius: 0.35rem;
  background: rgb(255 255 255 / 92%);
  color: var(--muted);
  font-size: 0.86rem;
  transform: translateX(-50%);
}

.figure-error {
  color: var(--danger);
}

.synopsis-figure figcaption {
  margin-top: 0.45rem;
  color: var(--muted);
  font-size: 0.86rem;
}

.site-footer {
  display: flex;
  justify-content: space-between;
  gap: 1rem;
  padding-block: 1.5rem 2.5rem;
  border-top: 1px solid var(--line);
  color: var(--muted);
  font-size: 0.85rem;
}

.site-footer p {
  margin: 0;
}

.visually-hidden {
  position: absolute !important;
  width: 1px !important;
  height: 1px !important;
  padding: 0 !important;
  margin: -1px !important;
  overflow: hidden !important;
  clip: rect(0, 0, 0, 0) !important;
  white-space: nowrap !important;
  border: 0 !important;
}

[hidden] {
  display: none !important;
}

@keyframes loading-sheen {
  to {
    background-position: -220% 0;
  }
}

@media (max-width: 700px) {
  .site-header__inner,
  .species-toolbar__inner,
  main,
  .site-footer {
    width: min(calc(100% - 1.25rem), var(--content-width));
  }

  .species-toolbar__inner,
  .species-header,
  .metadata-panel,
  .section-heading,
  .site-footer {
    align-items: stretch;
    flex-direction: column;
  }

  .species-toolbar__inner {
    gap: 0.55rem;
  }

  .species-picker {
    flex-basis: auto;
  }

  .species-nav {
    display: grid;
    grid-template-columns: 1fr 1fr;
  }

  .species-header {
    gap: 1.25rem;
  }

  .status-badges,
  .external-links {
    justify-content: flex-start;
  }

  .status-badges {
    flex-basis: auto;
  }

  .metadata-panel {
    gap: 1rem;
  }

  .section-heading > p {
    max-width: 32rem;
  }
}

@media (prefers-reduced-motion: reduce) {
  html {
    scroll-behavior: auto;
  }

  .figure-frame.is-loading {
    animation: none;
  }
}